Vlachos E. (2020) A response to Dubois (2020; Zoosystema 42 (23): 475-482). Zoosystema 42 (25): 515-517

Fernández, M. S., Vlachos, E., Buono, M. R., Alzugaray, L., Campos, L., Sterli, J., Herrera, Y., & Paolucci, F. (2020). Fingers zipped up or baby mittens? Two main tetrapod strategies to return to the sea. Biology Letters16(8), 20200281.

Vlachos, E. (2020) The turtles from ‘Gilmore’s slab’: evolution and diversity of the extinct turtle Echmatemys (Testudines: Testudinoidea: Geoemydidae) from the central United States. Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2020.1754938

Vlachos, E., Pérez-García, Roussiakis, S., Georgalis, G., Kear, B. (2020). Late Miocene tortoises from Samos island in Greece: implications for European Neogene testudinid systematics and distributions. Journal of Vertebrate Paleontology, e1722950. doi.org/10.1080/02724634.2020.1722950

Vlachos, E., Georgalis, G., Roussiakis, S., Böhme, M., Theodorou, G. (2020). The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of South Balkans. Journal of Vertebrate Paleontology, e1711520. doi.org/10.1080/02724634.2019.1711520

Pérez-García, A., Vlachos, E. , & Murelaga, X. (2020). A large testudinid tortoise with African affinities in the post-Messinian (Lower Pliocene) record of southeastern Spain. Palaeontology, doi: 10.1111/pala.12468


Vlachos, E. (2019). Introducing a new tool to navigate, understand and use International Codes of Nomenclature. PeerJ. doi 10.7717/peerj.8127

Vlachos, E. (2019). On the nomenclature of the largest tortoise that ever lived: Megalochelys sivalensis Falconer & Cautley, 1837 vs. Colossochelys atlas Falconer & Cautley, 1844 (Reptilia, Testudinidae). Bulletin of Zoological Nomenclature, 76: 162–164.

Vlachos, E., Sterli, J., Vasileiadou, K. & Syrides, G. (2019). A new species of Mauremys (Testudines, Geoemydidae) from the late Miocene – Pliocene of Central Macedonia (northern Greece) with exceptionally wide vertebral scutes. Papers in Palaeontology, doi.org/10.1002/spp2.1235.


Vlachos E., Bacharidis, N. (2018). A trionychid turtle from the late Aquitanian (early Miocene) of Neochori (Grevena, Western Macedonia, Greece). Bollettino della Societa Paleontologica Italiana, 57(3): 255-257

Vlachos, E., Randolfe, E., Sterli, J. & Leardi, J. M. (2018). Changes in the Diversity of Turtles (Testudinata) in South America from the Late Triassic to the Present. Ameghiniana, doi: 10.5710/AMGH.18.09.2018.3226

de la Fuente, M., Zacarías, G. & Vlachos, E. (2018). A review of the fossil record of the South American turtles of the clade Testudinoidea. Bulletin of the Peabody Museum of Natural History, 59(2):269-286

Athanassiou, A., Michailidis, D., Vlachos, E., Tourloukis, V., Thompson, N., & Harvati, K. (2018). Pleistocene vertebrates from the Kyparíssia lignite mine, Megalopolis Basin, S. Greece: Testudines, Aves, Suiformes. Quaternary International. doi.org/10.1016/j.quaint.2018.06.030

Vlachos, E., Tsoukala, E., Crégut-Bonnoure, E., Guérin, C., & Mol, D. (2018). The Paradise Lost of Milia (Grevena, Greece; Late Pliocene, Early Villafranchian, MN15/MN16a): Faunal Composition and Diversity. Quaternary1(2): 1–13.

Vlachos, E., Sterli, J. & de la Fuente, M. (2018). On the validity of Testudinites sellovii Weiss, 1830 from the Pleistocene of Paso del Catalán (Uruguay, South America). Ameghiniana, Paleontological Note, 55(3): 339–342.

Vlachos, E. (2018). A review of the fossil record of the North American turtles of the clade Pan-Testudinoidea. Bulletin of the Yale Peabody Museum, 59(1): 3–94.


Vlachos, E. & Rabi, M. (2017). Total evidence analysis and body size evolution of extant and extinct tortoises (Testudines: Cryptodira: Pan-Testudinidae). Cladistics. 1–32. 10.1111/cla.12227

Pérez-García, A., Vlachos, E. & Arribas, A. (2017). The last giant continental tortoise of Europe: a survivor in the Spanish Pleistocene site of Fonelas P-1. Palaeogeography, Palaeoclimatology, Palaeoecology, 47: 30–39.


Georgalis, G., Villa, A., Vlachos, E., Delfino, M. (2016). Fossil amphibians and reptiles from Plakias, Crete: a glimpse into the earliest late Miocene herpetofaunas of southeastern Europe. Geobios, 49: 433–444.

Pérez-García, A., Vlachos, E. & Mocho, P. (2016). The westernmost records of large European tortoises: the presence of Titanochelon (Testudinidae) in the Miocene of Portugal. Historical Biology. 29(6): 854–861.

Vlachos, E. & Delfino, M. (2016). Food for thought: sub-fossil and fossil chelonian remains from Franchthi Cave and Megalopolis confirm a Glacial refuge for Emys orbicularis in Peloponnesus (S. Greece). Quaternary Science Reviews150, 158-171. 10.1016/j.quascirev.2016.08.027.

Zougrou, IM., Katsikini, M., Pinakidou, F., Brzhezinskaya, M., Papadopoulou, L., Vlachos, E., Tsoukala, E. , Paloura, E. (2016). Characterization of fossil remains using XRF, XPS and XAFS spectroscopies. Journal of Physics: Conference Series, 712:012090, doi:10.1088/1742-6596/712/1/012090.

Vlachos, E. & Tsoukala, E. (2016). The diverse fossil chelonians from Milia (Late Pliocene, Grevena, Greece) with a new species of Testudo Linnaeus, 1758 (Testudines: Testudinidae). Papers in Palaeontology, Vol 2, Part 1: 72-86.


Vlachos, E. (2015). The fossil chelonians of Greece. Taxonomy – Stratigraphy – Palaeoecology - Evolution. PhD Thesis, School of Geology, Aristotle University of Thessaloniki, Greece

Vlachos, E. (2015). Forming taxon names from Greek words. Bionomina, 9: 1-26 pp.

Vlachos, E., Cerda, I. & Tsoukala, E. (2015). The first record of a soft–shelled turtle (Testudines: Pan-Trionychidae) from southern Balkans (Pliocene, Gefira, N. Greece) and new information from bone histology. The Science of Nature – Naturwissenschaften, 102, article 45, 9 pp.

Vlachos, E., Delfino, M. & Kotsakis, T. (2015). The chelonians from the latest Miocene-earliest Pliocene locality of Allatini and Pylea (east Thessaloniki, Macedonia, Greece). Comptes Rendus Palevol, 14(3): 187-205.


Vlachos, E. & Tsoukala, E. (2014). Testudo cf. graeca from the new Late Miocene locality of Platania (Drama basin, N. Greece) and a reappraisal of previously published specimens. Bulletin of the Geological Society of Greece, XLVIII: 27-40.

Pérez-García, A. & Vlachos, E. (2014). New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record. Zoological Journal of the Linnean Society, 172(3): 653-719.

Vlachos, E., Tsoukala, E. & Corsini, J. (2014). Cheirogaster bacharidisi sp. nov., a new species of a giant tortoise from the Pliocene of Thessaloniki (Macedonia, Greece). Journal of Vertebrate Paleontology, 34(3), 560-575.


Tsoukala, E., Mol, D., Pappa, S., Vlachos, E., Logchem, W., Vaxevanopoulos, M., Reumer, J. (2011). Elephas antiquus in Greece: New Finds and a reappraisal of older material (Mammalia, Proboscidea, Elephantidae). Quaternary International, 245: 339-349.


Pennos, C., Vaxevanopoulos, M., Pechlivanidou, S., Vlachos, E., Vavliakis, E. (2006). Determination of the flow velocity in cave conduits based on the study of scallops - „Eptamiloi‟ cave system, Macedonia, Greece. Proceedings of the 8th Hellenic Geographic Conference of the Hellenic Geographical Society, pp. 454 – 462.