logo conicet

Assistant Researcher

This email address is being protected from spambots. You need JavaScript enabled to view it.

 yatache 01 Twitter | Research Gate



I have completed my primary studies on the School of Geology in Aristotle University of Thessaloniki, Greece and soon I continued in the Master’s program on Tectonics & Stratigraphy. My primary focus is the study of fossil turtles, and my main field of research deals with the anatomy and evolution of terrestrial tortoises (Testudinidae). Testudinids form a clade with an almost worldwide distribution and an excellent, but not analyzed in detail fossil record. In my doctoral thesis I studied and described the entire fossil chelonian fauna of Greece, that is the most diverse and best preserved in the east Mediterranean. My research is expanded on the evolution of the large testudinid record of Europe.

This work has led me to pursue further research in Patagonia as a postdoctoral fellow in CONICET.

During my postdoctoral research in MEF, I focused on the origin and evolution of the extant and extinct testudinids of South America (clade Chelonoidis). These amazing turtles show an excellent fossil record, which I currently revise, and compare with similar forms in a global scale.

Now, my research focuses on the changes that happened in the biodiversity of turtles during their evolutionary history, and to discover previously unknown extinction events.

CV (March 2021)

2018ongoing – AGENCIA MinCyT PICT 2018 1848 “Origen, Extinción y Patrones de Diversificación de los Meiolaniformes y Pan-Quélidos (Testudinata) durante el intervalo Cretácico-Paleógeno en el sur de Sudamérica”. [Member of the Research Group]

20172019 – AGENCIA MinCyT PICT 2016 B 2334 “Análisis Filogenético de las tortugas terrestres (Testudines: Testudinidae) y la Historia Evolutiva de los testudínidos de Sudamérica”. [Lead Researcher]

2017ongoing – Editing of the volume “The Fossil Vertebrates of Greece”, to be published by Springer Nature in 2019. This is a volume that summarizes the entire fossil record of Greek vertebrates.

2016 – ongoing – Contributor to the PaleoBiology DataBase, focusing on the records of Testudinata (turtles).

20162018 – National Geographic Society / Waitt Grant W435-16 | "The Gulf of Titanic Turtles: excavation and preservation of new fossil tortoises from the Neogene of Northern Greece."

20142015 – IKY-DAAD, “Phylogenetic relationships and evolutionary history of the European Giant tortoises” in cooperation with the University of Tübingen, Germany.


Vlachos, E. (2021). A modularity analysis helps improving the structure of the International Code of Zoological Nomenclature. PeerJ9, e10815.


Vlachos E. (2020) A response to Dubois (2020; Zoosystema 42 (23): 475-482). Zoosystema 42 (25): 515-517

Fernández, M. S., Vlachos, E., Buono, M. R., Alzugaray, L., Campos, L., Sterli, J., Herrera, Y., & Paolucci, F. (2020). Fingers zipped up or baby mittens? Two main tetrapod strategies to return to the sea. Biology Letters16(8), 20200281.

Vlachos, E. (2020) The turtles from ‘Gilmore’s slab’: evolution and diversity of the extinct turtle Echmatemys (Testudines: Testudinoidea: Geoemydidae) from the central United States. Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2020.1754938

Vlachos, E., Pérez-García, Roussiakis, S., Georgalis, G., Kear, B. (2020). Late Miocene tortoises from Samos island in Greece: implications for European Neogene testudinid systematics and distributions. Journal of Vertebrate Paleontology, e1722950. doi.org/10.1080/02724634.2020.1722950

Vlachos, E., Georgalis, G., Roussiakis, S., Böhme, M., Theodorou, G. (2020). The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of South Balkans. Journal of Vertebrate Paleontology, e1711520. doi.org/10.1080/02724634.2019.1711520

Pérez-García, A., Vlachos, E. , & Murelaga, X. (2020). A large testudinid tortoise with African affinities in the post-Messinian (Lower Pliocene) record of southeastern Spain. Palaeontology, doi: 10.1111/pala.12468


Vlachos, E. (2019). Introducing a new tool to navigate, understand and use International Codes of Nomenclature. PeerJ. doi 10.7717/peerj.8127

Vlachos, E. (2019). On the nomenclature of the largest tortoise that ever lived: Megalochelys sivalensis Falconer & Cautley, 1837 vs. Colossochelys atlas Falconer & Cautley, 1844 (Reptilia, Testudinidae). Bulletin of Zoological Nomenclature, 76: 162–164.

Vlachos, E., Sterli, J., Vasileiadou, K. & Syrides, G. (2019). A new species of Mauremys (Testudines, Geoemydidae) from the late Miocene – Pliocene of Central Macedonia (northern Greece) with exceptionally wide vertebral scutes. Papers in Palaeontology, doi.org/10.1002/spp2.1235.


Vlachos E., Bacharidis, N. (2018). A trionychid turtle from the late Aquitanian (early Miocene) of Neochori (Grevena, Western Macedonia, Greece). Bollettino della Societa Paleontologica Italiana, 57(3): 255-257

Vlachos, E., Randolfe, E., Sterli, J. & Leardi, J. M. (2018). Changes in the Diversity of Turtles (Testudinata) in South America from the Late Triassic to the Present. Ameghiniana, doi: 10.5710/AMGH.18.09.2018.3226

de la Fuente, M., Zacarías, G. & Vlachos, E. (2018). A review of the fossil record of the South American turtles of the clade Testudinoidea. Bulletin of the Peabody Museum of Natural History, 59(2):269-286

Athanassiou, A., Michailidis, D., Vlachos, E., Tourloukis, V., Thompson, N., & Harvati, K. (2018). Pleistocene vertebrates from the Kyparíssia lignite mine, Megalopolis Basin, S. Greece: Testudines, Aves, Suiformes. Quaternary International. doi.org/10.1016/j.quaint.2018.06.030

Vlachos, E., Tsoukala, E., Crégut-Bonnoure, E., Guérin, C., & Mol, D. (2018). The Paradise Lost of Milia (Grevena, Greece; Late Pliocene, Early Villafranchian, MN15/MN16a): Faunal Composition and Diversity. Quaternary1(2): 1–13.

Vlachos, E., Sterli, J. & de la Fuente, M. (2018). On the validity of Testudinites sellovii Weiss, 1830 from the Pleistocene of Paso del Catalán (Uruguay, South America). Ameghiniana, Paleontological Note, 55(3): 339–342.

Vlachos, E. (2018). A review of the fossil record of the North American turtles of the clade Pan-Testudinoidea. Bulletin of the Yale Peabody Museum, 59(1): 3–94.


Vlachos, E. & Rabi, M. (2017). Total evidence analysis and body size evolution of extant and extinct tortoises (Testudines: Cryptodira: Pan-Testudinidae). Cladistics. 1–32. 10.1111/cla.12227

Pérez-García, A., Vlachos, E. & Arribas, A. (2017). The last giant continental tortoise of Europe: a survivor in the Spanish Pleistocene site of Fonelas P-1. Palaeogeography, Palaeoclimatology, Palaeoecology, 47: 30–39.


Georgalis, G., Villa, A., Vlachos, E., Delfino, M. (2016). Fossil amphibians and reptiles from Plakias, Crete: a glimpse into the earliest late Miocene herpetofaunas of southeastern Europe. Geobios, 49: 433–444.

Pérez-García, A., Vlachos, E. & Mocho, P. (2016). The westernmost records of large European tortoises: the presence of Titanochelon (Testudinidae) in the Miocene of Portugal. Historical Biology. 29(6): 854–861.

Vlachos, E. & Delfino, M. (2016). Food for thought: sub-fossil and fossil chelonian remains from Franchthi Cave and Megalopolis confirm a Glacial refuge for Emys orbicularis in Peloponnesus (S. Greece). Quaternary Science Reviews150, 158-171. 10.1016/j.quascirev.2016.08.027.

Zougrou, IM., Katsikini, M., Pinakidou, F., Brzhezinskaya, M., Papadopoulou, L., Vlachos, E., Tsoukala, E. , Paloura, E. (2016). Characterization of fossil remains using XRF, XPS and XAFS spectroscopies. Journal of Physics: Conference Series, 712:012090, doi:10.1088/1742-6596/712/1/012090.

Vlachos, E. & Tsoukala, E. (2016). The diverse fossil chelonians from Milia (Late Pliocene, Grevena, Greece) with a new species of Testudo Linnaeus, 1758 (Testudines: Testudinidae). Papers in Palaeontology, Vol 2, Part 1: 72-86.


Vlachos, E. (2015). The fossil chelonians of Greece. Taxonomy – Stratigraphy – Palaeoecology - Evolution. PhD Thesis, School of Geology, Aristotle University of Thessaloniki, Greece

Vlachos, E. (2015). Forming taxon names from Greek words. Bionomina, 9: 1-26 pp.

Vlachos, E., Cerda, I. & Tsoukala, E. (2015). The first record of a soft–shelled turtle (Testudines: Pan-Trionychidae) from southern Balkans (Pliocene, Gefira, N. Greece) and new information from bone histology. The Science of Nature – Naturwissenschaften, 102, article 45, 9 pp.

Vlachos, E., Delfino, M. & Kotsakis, T. (2015). The chelonians from the latest Miocene-earliest Pliocene locality of Allatini and Pylea (east Thessaloniki, Macedonia, Greece). Comptes Rendus Palevol, 14(3): 187-205.


Vlachos, E. & Tsoukala, E. (2014). Testudo cf. graeca from the new Late Miocene locality of Platania (Drama basin, N. Greece) and a reappraisal of previously published specimens. Bulletin of the Geological Society of Greece, XLVIII: 27-40.

Pérez-García, A. & Vlachos, E. (2014). New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record. Zoological Journal of the Linnean Society, 172(3): 653-719.

Vlachos, E., Tsoukala, E. & Corsini, J. (2014). Cheirogaster bacharidisi sp. nov., a new species of a giant tortoise from the Pliocene of Thessaloniki (Macedonia, Greece). Journal of Vertebrate Paleontology, 34(3), 560-575.


Tsoukala, E., Mol, D., Pappa, S., Vlachos, E., Logchem, W., Vaxevanopoulos, M., Reumer, J. (2011). Elephas antiquus in Greece: New Finds and a reappraisal of older material (Mammalia, Proboscidea, Elephantidae). Quaternary International, 245: 339-349.


Pennos, C., Vaxevanopoulos, M., Pechlivanidou, S., Vlachos, E., Vavliakis, E. (2006). Determination of the flow velocity in cave conduits based on the study of scallops - „Eptamiloi‟ cave system, Macedonia, Greece. Proceedings of the 8th Hellenic Geographic Conference of the Hellenic Geographical Society, pp. 454 – 462.

Networks of the International Codes of Nomenclature

Fig 2 Code Structure


Vlachos E. 2019. Introducing a new tool to navigate, understand and use International Codes of Nomenclature. PeerJ 7:e8127 http://doi.org/10.7717/peerj.8127

Vlachos E. 2021. A modularity analysis helps improving the structure of the International Code of Zoological Nomenclature. PeerJ 9:e10815 https://peerj.com/articles/10815/



Neticon zoologicon

The Network of the International Code of Zoological Nomenclature

Disclaimer: The International Code of Zoological Nomenclature is published by the International Trust for Zoological Nomenclature 1999. The text of the Code is reproduced herein with the written consent of the publisher.

The Code is available here


Current version | version 1.0 [publication date: 24 October 2019]

The version 1.0 of the Neticon zoologicon contains the 4th edition of the ICZN (1999), including the 2012 Amendment on electronic publication, and Declarations 44, and 45.

Neticon_zoologiconv1.0 Html (Online) Version

(a new tab will open with the online html version of the Neticon)

Neticon_zoologiconv1.0 Html (Offline) Version

(a zipped file will be downloaded, containing the html version of the Neticon.

Unzip the folder, and open the file index.html with any browser except Google Chrome)

Neticon_zoologiconv1.0 Gephi Version (original and editable)

(a zipped file will be downloaded, containing the original network in Gephi format)



Neticon phylogeneticon

The Network of the PhyloCode

The beta version of the Neticon phylogeneticon contains the version 6 of the PhyloCode (2020), written by Philip D. Cantino and Kevin de Queiroz, published by CRC Press of the Taylor & Francis Group. I have permission to reproduce this text herein.

For more information on the PhyloCode, please check here. Buy the PhyloCode here, and the Phylonyms here.


Current version | beta version [publication date: 18 November 2020]

Neticon_phylogeneticonbeta Html (Online) Version

(a new tab will open with the online html version of the Neticon)

This beta html version is designed to accompany the poster presentation in the 2020 PalAss Virtual Annual Meeting.





A series of videos explaining some of the functions and abilities of the Neticon

Neticon_zoologicon Tutorial 1: Basic Introduction to the Online Version. 

Neticon_zoologicon Tutorial 2: Basic Introduction to the Gephi Version.

Neticon_zoologicon Tutorial 3: How to use the Ego Network filter, to create subsets of the Code.


> Questions that you might want to ask <

What is it?

This is a network analysis of the Code. All the elements of the Code (Articles, Recommendations, Examples, Glossary Items) are interconnected according to their direct and indirect connections. After the network is created, some structural layouts are applied to depict its structure.

How can I use it?

You could click on the links above for the different versions. The first two links (marked with Online and Offline) contains the exported network with the Force Atlas 2 layout. The user can navigate through the network, but cannot make any changes. The last link (marked with Gephi) is the original network to be opened with Gephi. There the user can enjoy the full capabilities of the network.

Please read the manuscript and especially the supplements for further information.

What shall I do if I discover any problems, malfunctions, errors, etc.?

Please report them to me by email (This email address is being protected from spambots. You need JavaScript enabled to view it.). I will do my best to fix them.

Can I edit the network?

Of course!

I would encourage users to try to further develop this effort, correct mistakes and try different things and layouts. As I am planning to regularly release updates, if you do so, however, please inform me. In case you have made a significant update or change, your version could be hosted here.

Also, I am open to any kind of collaboration and further development. So please do not hesitate to contact me!

If I have questions?

If there are software-related please refer to the appropriate sources. The Gephi and Sigma communities are quite open and active, and host wonderful tutorials and forums.

For anything else, please do not hesitate to email me and I will do my best to respond.


May the Code be with you…


Evan Vlachos,

CONICET and Museo Paleontológico Egidio Feruglio,

Av. Fontana 140, 9100, Trelew, Chubut, Argentina

This email address is being protected from spambots. You need JavaScript enabled to view it.

Last Update: February 23, 2021

Flag Counter