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Postdoctoral Scholar

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 yatache 01


I have completed my primary studies on the School of Geology in Aristotle University of Thessaloniki, Greece and soon I continued in the Master’s program on Tectonics & Stratigraphy. My primary focus is the study of fossil turtles, and my main field of research deals with the anatomy and evolution of terrestrial tortoises (Testudinidae). Testudinids form a clade with an almost worldwide distribution and an excellent, but not analyzed in detail fossil record. In my doctoral thesis I studied and described the entire fossil chelonian fauna of Greece, that is the most diverse and best preserved in the east Mediterranean. My research is expanded on the evolution of the large testudinid record of Europe.

This work has led me to pursue further research in Patagonia as a postdoctoral fellow in CONICET.

During my postdoctoral research in MEF, I focused on the origin and evolution of the extant and extinct testudinids of South America (clade Chelonoidis). These amazing turtles show an excellent fossil record, which I currently revise, and compare with similar forms in a global scale.

Now, my research focuses on the changes that happened in the biodiversity of turtles during their evolutionary history, and to discover previously unknown extinction events.

CV (March 2018)

2017ongoing – AGENCIA MinCyT PICT B 2334 “Análisis Filogenético de las tortugas terrestres (Testudines: Testudinidae) y la Historia Evolutiva de los testudínidos de Sudamérica”.

2017ongoing – Editing of the volume “The Fossil Vertebrates of Greece”, to be published by Springer Nature in 2019. This is a volume that summarizes the entire fossil record of Greek vertebrates.

2016 – ongoing – Contributor to the PaleoBiology DataBase, focusing on the records of Testudinata (turtles).

20162018 – National Geographic Society / Waitt Grant W435-16 | "The Gulf of Titanic Turtles: excavation and preservation of new fossil tortoises from the Neogene of Northern Greece."

20142015 – IKY-DAAD, “Phylogenetic relationships and evolutionary history of the European Giant tortoises” in cooperation with the University of Tübingen, Germany.

Vlachos, E., Sterli, J., Vasileiadou, K. & Syrides, G. (2019). A new species of Mauremys (Testudines, Geoemydidae) from the late Miocene – Pliocene of Central Macedonia (northern Greece) with exceptionally wide vertebral scutes. Papers in Palaeontology, doi.org/10.1002/spp2.1235.

Vlachos E., Bacharidis, N. (2018). A trionychid turtle from the late Aquitanian (early Miocene) of Neochori (Grevena, Western Macedonia, Greece). Bollettino della Societa Paleontologica Italiana, 57(3): 255-257

Vlachos, E., Randolfe, E., Sterli, J. & Leardi, J. M. (2018). Changes in the Diversity of Turtles (Testudinata) in South America from the Late Triassic to the Present. Ameghiniana, doi: 10.5710/AMGH.18.09.2018.3226

de la Fuente, M., Zacarías, G. & Vlachos, E. (2018). A review of the fossil record of the South American turtles of the clade Testudinoidea. Bulletin of the Peabody Museum of Natural History, 59(2):269-286

Athanassiou, A., Michailidis, D., Vlachos, E., Tourloukis, V., Thompson, N., & Harvati, K. (in press). Pleistocene vertebrates from the Kyparíssia lignite mine, Megalopolis Basin, S. Greece: Testudines, Aves, Suiformes. Quaternary International. doi.org/10.1016/j.quaint.2018.06.030

Vlachos, E., Tsoukala, E., Crégut-Bonnoure, E., Guérin, C., & Mol, D. (2018). The Paradise Lost of Milia (Grevena, Greece; Late Pliocene, Early Villafranchian, MN15/MN16a): Faunal Composition and Diversity. Quaternary1(2): 1–13.

Vlachos, E., Sterli, J. & de la Fuente, M. (2018). On the validity of Testudinites sellovii Weiss, 1830 from the Pleistocene of Paso del Catalán (Uruguay, South America). Ameghiniana, Paleontological Note, 55(3): 339–342.

Vlachos, E. (2018). A review of the fossil record of the North American turtles of the clade Pan-Testudinoidea. Bulletin of the Yale Peabody Museum, 59(1): 3–94.

Vlachos, E. & Rabi, M. (2017). Total evidence analysis and body size evolution of extant and extinct tortoises (Testudines: Cryptodira: Pan-Testudinidae). Cladistics. 1–32. 10.1111/cla.12227

Pérez-García, A., Vlachos, E. & Arribas, A. (2017). The last giant continental tortoise of Europe: a survivor in the Spanish Pleistocene site of Fonelas P-1. Palaeogeography, Palaeoclimatology, Palaeoecology, 47: 30–39.

Georgalis, G., Villa, A., Vlachos, E., Delfino, M. (2016). Fossil amphibians and reptiles from Plakias, Crete: a glimpse into the earliest late Miocene herpetofaunas of southeastern Europe. Geobios, 49: 433–444.

Pérez-García, A., Vlachos, E. & Mocho, P. (2016). The westernmost records of large European tortoises: the presence of Titanochelon (Testudinidae) in the Miocene of Portugal. Historical Biology. 29(6): 854–861.

Vlachos, E. & Delfino, M. (2016). Food for thought: sub-fossil and fossil chelonian remains from Franchthi Cave and Megalopolis confirm a Glacial refuge for Emys orbicularis in Peloponnesus (S. Greece). Quaternary Science Reviews150, 158-171. 10.1016/j.quascirev.2016.08.027.

Zougrou, IM., Katsikini, M., Pinakidou, F., Brzhezinskaya, M., Papadopoulou, L., Vlachos, E., Tsoukala, E. , Paloura, E. (2016). Characterization of fossil remains using XRF, XPS and XAFS spectroscopies. Journal of Physics: Conference Series, 712:012090, doi:10.1088/1742-6596/712/1/012090.

Vlachos, E. & Tsoukala, E. (2016). The diverse fossil chelonians from Milia (Late Pliocene, Grevena, Greece) with a new species of Testudo Linnaeus, 1758 (Testudines: Testudinidae). Papers in Palaeontology, Vol 2, Part 1: 72-86.

Vlachos, E. (2015). Forming taxon names from Greek words. Bionomina, 9: 1-26 pp.

Vlachos, E., Cerda, I. & Tsoukala, E. (2015). The first record of a soft–shelled turtle (Testudines: Pan-Trionychidae) from southern Balkans (Pliocene, Gefira, N. Greece) and new information from bone histology. The Science of Nature – Naturwissenschaften, 102, article 45, 9 pp.

Vlachos, E., Delfino, M. & Kotsakis, T. (2015). The chelonians from the latest Miocene-earliest Pliocene locality of Allatini and Pylea (east Thessaloniki, Macedonia, Greece). Comptes Rendus Palevol, 14(3): 187-205.

Vlachos, E. & Tsoukala, E. (2014). Testudo cf. graeca from the new Late Miocene locality of Platania (Drama basin, N. Greece) and a reappraisal of previously published specimens. Bulletin of the Geological Society of Greece, XLVIII: 27-40.

Pérez-García, A. & Vlachos, E. (2014). New generic proposal for the European Neogene large testudinids (Cryptodira) and the first phylogenetic hypothesis for the medium and large representatives of the European Cenozoic record. Zoological Journal of the Linnean Society, 172(3): 653-719.

Vlachos, E., Tsoukala, E. & Corsini, J. (2014). Cheirogaster bacharidisi sp. nov., a new species of a giant tortoise from the Pliocene of Thessaloniki (Macedonia, Greece). Journal of Vertebrate Paleontology, 34(3), 560-575.

Tsoukala, E., Mol, D., Pappa, S., Vlachos, E., Logchem, W., Vaxevanopoulos, M., Reumer, J. (2011). Elephas antiquus in Greece: New Finds and a reappraisal of older material (Mammalia, Proboscidea, Elephantidae). Quaternary International, 245: 339-349.

Pennos, C., Vaxevanopoulos, M., Pechlivanidou, S., Vlachos, E., Vavliakis, E. (2006). Determination of the flow velocity in cave conduits based on the study of scallops - „Eptamiloi‟ cave system, Macedonia, Greece. Proceedings of the 8th Hellenic Geographic Conference of the Hellenic Geographical Society, pp. 454 – 462.

under construction...

Scientific Graphic Design has always been a passion for me. The ability to transform scientific information into a design that will inform but will also capture the interest of professionals and non-professionals is a challenge for me. What started as a hobby, has now become an obsession. Throughout my career, I have received professional training in graphic design. I was able to apply this knowledge and my ideas in several cases, including paleontological exhibitions, books, posters, and figures in scientific publications. A selection of my favorite designs is given below.

Paleontological Exhibitions


2018 | New Paleontological Exhibition of Milia, Grevena, Greece, with the longest tusks in the world!  



Networks of the International Codes of Nomenclature

DisclaimerThe International Code of Zoological Nomenclature is published by the International Trust for Zoological Nomenclature 1999. The text of the Code is reproduced herein with the written consent of the publisher.

The Code is available at: http://www.iczn.org/iczn/index.jsp



Neticon zoologicon

The Network of the International Code of Zoological Nomenclature


Current version | Beta version [publication date: June 2019]

The beta version of the Neticon contains the 4th edition of the ICZN (1999), including the 2012 Amendment on electronic publication, and Declarations 44, and 45.

Neticon_zoologiconbeta Html (Online) Version    [link active]

(a new tab will open with the online html version of the Neticon)

Neticon_zoologiconbeta Html (Offline) Version     [link NOT active]

(a zipped file will be downloaded, containing the html version of the Neticon.

Unzip the folder, and open the file index.html with any browser except Google Chrome)

Neticon_zoologiconbeta Gephi Version     [link NOT active]

(a zipped file will be downloaded, containing the original network in Gephi format)





A series of videos explaining some of the functions and abilities of the Neticon

Neticon_zoologicon Tutorial 1: Basic Introduction to the Online Version. 

Neticon_zoologicon Tutorial 2: Basic Introduction to the Gephi Version.

Neticon_zoologicon Tutorial 3: How to use the Ego Network filter, to create subsets of the Code.


> Frequently Asked Questions <

What is it?

This is a network analysis of the Code. All the elements of the Code (Articles, Recommendations, Examples, Glossary Items) are interconnected according to their direct and indirect connections. After the network is created, some structural layouts are applied to depict its structure.

How can I use it?

You could click on the links above for the different versions. The first link (marked with Online) contains the exported network with the Force Atlas 2 layout. The user can navigate through the network, but cannot make any changes. The last link (marked with Gephi) is the original network to be opened with Gephi. There the user can enjoy the full capabilities of the network. Please read the manuscript for further information [link].

What shall I do if I discover any problems, malfunctions, errors, etc.?

Please report them to me by email (This email address is being protected from spambots. You need JavaScript enabled to view it.). I will do my best to fix them.

Can I edit the network?

Of course! I would encourage users to try to further develop this effort, correct mistakes and try different things and layouts. As I am planning to regularly release updates, if you do so, however, please inform me. In case you have made a significant update or change, your version could be hosted here.

If I have questions?

If there are software-related please refer to the appropriate source.

For anything else, please do not hesitate to email me and I will do my best to respond.


May the Code be with you…


Evan Vlachos,

CONICET and Museo Paleontológico Egidio Feruglio,

Av. Fontana 140, 9100, Trelew, Chubut, Argentina

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The networks herein are shared with a Attribution-NonCommercial-ShareAlike 3.0 Unported (CC BY-NC-SA 3.0).

You are free to:

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Last Update: June 11, 2019

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